MAPK/ERK Signaling Pathway

The MAPK/ERK pathway is a critical signaling cascade that regulates cell proliferation, differentiation, and survival.

Overview

The MAPK/ERK pathway (also known as the RAS-RAF-MEK-ERK pathway) is one of the most well-studied signaling cascades in cell biology. It transmits signals from cell surface receptors to the nucleus, regulating gene expression involved in cell proliferation, differentiation, and survival.

Key Components

ComponentFunctionDysregulation in Disease
RAS (KRAS, NRAS, HRAS)Small GTPase that activates RAFMutated in ~30% of all cancers
RAF (ARAF, BRAF, CRAF)Serine/threonine kinaseBRAF V600E mutation in melanoma
MEK (MEK1, MEK2)Dual-specificity kinaseOveractive in various cancers
ERK (ERK1, ERK2)Proline-directed kinaseHyperactivated in many tumors

Signaling Mechanism

Upon growth factor binding to receptor tyrosine kinases (RTKs), RAS is activated through GDP-to-GTP exchange. Activated RAS recruits RAF to the membrane, which phosphorylates and activates MEK. MEK then phosphorylates ERK, which phosphorylates numerous cytoplasmic and nuclear targets.

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